传说森林住着邪恶的大魔王，为了帮助小精灵脱险，羊村守护者们再度出击勇闯四季城。不料，途中喜羊羊竟意外变成无法自控的“破影大王”，时而清醒，时而捣乱，让整个旅途笑料百出。羊狼们一路闯关进阶“勇者”能力，
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少女战士法尼娅遇到了名为“导师”的失忆少年，两人共同展开旅程，逐渐揭开敌人——“异生体”的秘密。与此同时也发现了“某个神秘势力”隐藏在这之后想要操控少女战士“斯露德”的人生。她们决定奋起反抗，净化土地
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